In the earliest cortical stages of visual processing, a scene is represented in different functional domains selective for specific image features. Maps of orientation and spatial frequency preference have been described in the primary visual cortex using simple sinusoidal grating stimuli. However, recent imaging experiments suggest that the maps of these two spatial parameters are not sufficient to describe patterns of activity generated in response to complex moving stimuli (Basole, et al. 2003). A model of cortical organization is presented in which temporal frequency tuning is superimposed on the maps of orientation and spatial frequency tuning previously measured in cat and ferret Area 17. The organization of temporal frequency tuning used in this model is based on measurements made in cat Area 17 that suggest temporal frequency preference and bandwidth vary in a subtle but systematic way across different spatial frequency domains (Zhang and Issa, SFN 2004). The combined map of these three filtering p...