Hearing and Communication Neuroscience (HCN) is an inter-disciplinary training program for graduate students, post-doctoral scholars, and physician-scientists at the University of Southern California. Cutting-edge research labs are investigating mechanisms of audition and vocal communication and are located in the Departments of Otolaryngology, Stem Cell Biology, Physiology and Biophysics, Neurobiology, Linguistics, Psychology, and Biomedical Engineering. Our training program brings together researchers spanning multiple disciplines, ranging from genetics and cell-molecular biology to cognitive, systems, and behavioral neuroscience. HCN labs are investigating mechanisms underlying audition and vocal communication at the level of individual molecules and cells, neural circuits, and cognitive constructs such as neural representations of memories, plans and actions. Our program is supported by an NIH training grant from the National Institute of Deafness and Communication Disorders. Graduate Students must be accepted by the Neuroscience Graduate Program at USC, and can apply to the HCN Program at the end of their first year once they have chosen a home lab; applications for graduate study can be made at https://ngp.usc.edu/. Postdoctoral Fellows and Physician-Scientists can initiate an application by contacting individual Faculty through our website at http://dornsife.usc.edu/hcn/.